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    1. Species/
    2. Echinoderms/
    3. Protoreaster nodosus

    Echinoderms · Oreasteridae

    Protoreaster nodosus

    Chocolate chip sea star

    A large, rigid sea star of western Pacific seagrass beds and sandy bottoms, studded with black conical 'horns' that earn it the name chocolate chip star. Hardy, but not reef safe: adults eat soft corals, sponges, clams, tubeworms and other stars.

    • intermediate level
    • Not reef safe
    Minimum tank
    500 L
    Adult size
    ≤ 30 cm
    Lifespan
    ≤ 17 years

    This sheet is still being written. Some sections are not filled in yet: they stay hidden while they are empty.

    On this sheet

    1. Water parameters
    2. Care and feeding
    3. Compatibility
    4. Description
    5. Conservation
    6. Classification
    7. Sources

    What it needs

    Water parameters

    • Temperature°C
      2230
      22.2–25.6 °C
    • SalinitySG
      1.021.03
      1.023–1.025 SG

    Related species

    Other echinoderms

    See all echinoderms →

    Blue linckia

    Linckia laevigata

    E

    Sand star

  1. pH
    7.68.6
    8.1–8.4
  2. Range recommended by the sheet

    Day to day

    Care and feeding

    • LevelIntermediate
    • FoodCarnivore
    • Tank≥ 500 L

    Care

    Not for a reef aquarium: adults eat soft corals, sponges, tubeworms, clams and other sea stars, and Quality Marine also lists anemones, gorgonians, stony corals, urchins and small unwary fish. Keep it in a large fish-only tank with live rock (FOWLR).

    Acclimation: it cannot tolerate rapid changes in water chemistry (oxygen, salinity, pH). Acclimate it by drip and never expose it to air while handling.

    Setup: a very large tank (Reeflex recommends at least 500 L) with open access to a mature sand bed several centimetres deep and plenty of open space.

    Tankmates and sensitivities: keep it with calm fish; triggerfish and pufferfish are its predators. It needs very good water quality and does not tolerate high nitrate, copper or most anti-parasite treatments.

    Water: aim for 22 to 25.5 °C, a specific gravity of 1.023 to 1.025, pH 8.1 to 8.4 and 8 to 12 dKH. Sources rate it anywhere from 'easy' (LiveAquaria) to 'difficult' (Quality Marine); Reeflex calls it 'average'.

    Feeding

    In the wild it feeds mostly on algal and microbial films, living and decaying seagrass blades and filamentous green algae, which it digests by everting its stomach onto the substrate. It is also an opportunistic carnivore: molluscs, sponges, soft corals, bryozoans, tubeworms, other stars; one was seen eating a sea urchin.

    In the aquarium, feed it chopped clam, shrimp and squid. It also hunts down the snails in the tank.

    Decor

    • Mature sand bed several centimetres deep, freely accessible
    • Large open space
    • Live rock

    Tank mates

    Compatibility

    Goes well with

    • ✓Large fish-only tank with live rock (FOWLR)
    • ✓Calm fish, too large to be prey

    Avoid

    • ✕Soft and stony corals, anemones, gorgonians, sponges: it eats or disturbs them
    • ✕Clams (Tridacna), snails, tubeworms, urchins and other sea stars: prey
    • ✕Small unwary fish
    • ✕Triggerfish and pufferfish: predators
    • ✕Copper and most anti-parasite treatments

    In detail

    Description

    Identification: five short, sturdy arms and a domed central disc; a row of dark conical horns on each arm and a cluster on the disc, which may wear blunt. Colour varies: cream, light tan, orange, red, brown, sometimes blue or yellow depending on origin; white underside, purple or pale pink tube feet. 15 cm on average, 30 cm according to SeaLifeBase and up to 40 cm according to other sources. The horns do not sting: the animal is harmless.

    In the wild it prefers sheltered sandy or muddy bottoms and the seagrass beds of calm lagoons, from the surface to about 30 m, rarely below 10 m; it can be left exposed at low tide. It gathers in aggregations (over 5 individuals per 10 m²) where food is plentiful. It mostly grazes algal and microbial films and seagrass blades, but also attacks large prey, including other stars such as Linckia laevigata. A 2022 revision limits its range to the western Pacific; records from the Indian Ocean probably refer to the sister species Protoreaster lincki.

    Reproduction and status: separate sexes, fertilisation in open water, planktonic larvae for a few weeks; mature at about 12 cm, possibly living up to 17 years. Heavily collected as a dried souvenir, it has collapsed locally (not seen on Guam since 1945). It has not been evaluated by the IUCN and is not listed by CITES.

    Characteristics

    • Black conical horns on the arms and disc; they do not sting.
    • 15 cm on average, 30 to 40 cm at most depending on the source.
    • Not reef safe: eats soft corals, sponges, clams, tubeworms and other stars.
    • Seagrass beds and sandy bottoms of calm lagoons, mostly shallower than 10 m.
    • May live up to 17 years.
    • Heavily collected as a dried souvenir.

    Distribution

    • Tropical western Pacific, from Thailand to Samoa and from Japan to New Caledonia (2022 revision)
    • Surface to about 30 m, mostly shallower than 10 m

    Status

    Conservation

    IUCN Red List

    Not evaluated (NE)

    WoRMS

    Classification

    Kingdom
    Animalia
    └
    Phylum
    Echinodermata
    └
    Class
    Asteroidea
    └
    Order
    Valvatida
    └
    Family
    Oreasteridae
    └
    Genus
    Protoreaster
    └
    Species
    Protoreaster nodosus

    AphiaID 213285 · See on WoRMS (opens in a new tab)

    Where this comes from

    Sources

    1. [1]WoRMSWoRMS — Protoreaster nodosus (Linnaeus, 1758) (AphiaID 213285) (opens in a new tab) · accessed on November 11, 2026
    2. [2]WebSeaLifeBase — Protoreaster nodosus, Horned sea star (species summary) (opens in a new tab) · accessed on November 11, 2026
    3. [3]WebGBIF — IUCN Red List category of Protoreaster nodosus (Not Evaluated) (opens in a new tab) · accessed on November 11, 2026
    4. [4]CITESSpecies+ — search for Protoreaster and Protoreaster nodosus (no CITES taxon concept) (opens in a new tab) · accessed on November 11, 2026
    5. [5]WebWikipedia (en) — Protoreaster nodosus (description, habitat, diet in aquariums, overharvesting) (opens in a new tab) · accessed on November 11, 2026
    6. [6]WebWikipédia (fr) — Protoreaster nodosus (size, ecology, reproduction, lifespan, predators, collection) (opens in a new tab) · accessed on November 11, 2026
    7. [7]WebWikipedia (de) — Knotiger Walzenseestern (diet) (opens in a new tab) · accessed on November 11, 2026
    8. [8]WebWikipedia (it) — Protoreaster nodosus (opens in a new tab) · accessed on November 11, 2026
    9. [9]WebLiveAquaria — Chocolate Chip Sea Star, Protoreaster nodosus (quick stats and overview) (opens in a new tab) · accessed on November 11, 2026
    10. [10]WebQuality Marine — Chocolate Chip Sea Star, sold as Protoreaster sp. (not reef safe; care level) (opens in a new tab) · accessed on November 11, 2026
    11. [11]WebReeflex — Protoreaster nodosus (aquarium difficulty: average; at least 500 L) (opens in a new tab) · accessed on November 11, 2026
    12. [12]PaperDucarme, F. (2022). How to assess the absence of a species? A revision of the geographical range of the horned sea star, Protoreaster nodosus. Frontiers of Biogeography 14(3) (read through fr.wikipedia) · DOI 10.21425/F5FBG56187 (opens in a new tab)
    13. [13]PaperScheibling, R.E. & Metaxas, A. (2008). Abundance, spatial distribution, and size structure of the sea star Protoreaster nodosus in Palau, with notes on feeding and reproduction. Bulletin of Marine Science 82(2): 221-235 (read through fr.wikipedia)
    14. [14]WebLiveAquaria — Acclimation Guides (why acclimate; floating and drip methods) (opens in a new tab) · accessed on November 11, 2026
    15. [15]WebLiveAquaria — Employ the Drip Method to Reduce Aquarium Acclimation Stress (opens in a new tab) · accessed on November 11, 2026
    UpdatedNov 11, 2026
    Versionv2
    Report an error on this sheet

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